Extinction Archives

Great Auk

Extinct | 1844

Pinguinus impennis

Charadriiformes - Alcidae - Pinguinus

The Great Auk, a flightless seabird, was once widespread in the waters and islands of the North Atlantic. It was the only flightless species among modern auk family. The Great Auk and the extant Razorbill are evolutionary relatives. They both belonged to a single lineage distributed across the Atlantic during ancient times, and by the early Pliocene, they had each adapted to different environmental niches1.

The Great Auk measured about 75 to 80 centimeters in length, making it the largest modern species in the auk family2. Males and females looked alike, with black backs, white bellies, and heads and necks tinged with brown. They had small heads and streamlined bodies. The Great Auk’s plumage and body shape were remarkably similar to those of penguins, despite being unrelated, a prime example of convergent evolution. It had a hard, long beak with white grooves arranged diagonally, and oval white patches in front of its eyes. During the winter molt, its throat feathers turned white and the eye patches became thinner and lighter3. The Great Auk’s wings were short and highly reduced, darker than its body, and brown in color. Its feet were webbed and black.

Although clumsy on land, the Great Auk was an adept swimmer and diver. It inhabited the high-latitude waters of the North Atlantic, spending most of the year at sea, diving to catch fish. The Great Auk only came ashore to breed in the summer. They did not build nests; instead, they laid eggs directly on rocks, typically at the base of coastal cliffs. The Great Auk had specific breeding requirements, usually preferring isolated North Atlantic islands far from the mainland, with rocky shores suitable for laying eggs and accessing the sea. During the breeding season, they were highly social, with each pair producing one egg per season.

Humans have hunted the Great Auk for a very long time; indigenous peoples hunted them over a hundred thousand years ago. However, their decline towards extinction truly began during the Age of Exploration. Increased human activity in the North Atlantic and systematic commercial slaughter expanded. The Great Auk was hunted extensively for its meat, oil, feathers, and eggs. As their population declined, collectors’ demand for their skins and eggs increased, leading to intensified hunting.

By the late 18th century, the Great Auk had disappeared from breeding sites on the North American side of the Atlantic, and those in European waters gradually vanished too. The last groups migrated to remote islands near Iceland, where they were eventually hunted to extinction by humans in the mid-19th century.

The last confirmed sighting of a Great Auk was on June 3, 1844, on Eldey Island, Iceland, where three men killed a pair of Great Auks 4 incubating an egg for a bounty5. Later reports indicated that four Great Auks were sighted in April 1848 in the waters off East Finnmark, Norway, with one being shot but its remains lost to the sea6. An unconfirmed sighting occurred on December 1852 on the Grand Banks of Newfoundland, Canada7.

museum
Royal Ontario Museum, own work, taken on 2023.12.9
Later learned that this is the only Great Auk specimen in Canada! This makes me feel both lucky and sad. See: 1, 2
This specimen was originally part of the collection of the renowned ornithologist John James Audubon (below is one of his paintings of the Great Auk). It changed hands several times before being purchased by the Royal Ontario Museum. For more background, see (Fuller, 1999).
The Great Auk was once abundant on Funk Island off the eastern coast of Newfoundland, Canada, which was their largest habitat in modern times. Ultimately, Canada was left with this single specimen, as most others were sold to European collectors. The word "left" is in quotes because this sole Canadian specimen actually came from Iceland.
Great Auk, drawn by John James Audubon (1785-1851), Public Domain
The taxidermied specimen of the last male Great Auk killed in 1844
Royal Belgian Institute of Natural Sciences, photographed by Thierry Hubin, sourced from (Thomas 2017) (CC BY 4.0).
It was only in modern times that scientists, through DNA sequencing, identified the skin of the last male Great Auk. The whereabouts of the skin of the last female Great Auk remain unknown.
Two Great Auk specimens.
Left: Natural History Museum of the University of Pisa. See here; Right: Natural History Museum, London. Source: Wikimedia Commons
Another specimen preserved at the Natural History Museum of Denmark, killed in 1823 in Eyrabakki, Iceland.
In the background on the second photo are the internal organs of the last pair of Great Auks killed in 1844, preserved in Copenhagen.
From the left: 1. Photograph by Maja Theodoraki, sourced from (Kjartansdóttir 2019) (CC BY-NC-ND.4.0) | 2. 3. Photographs by Michael B. H., Wikimedia Commons
Great Auk specimen at the Cincinnati Museum. The skin used for this taxidermy may have belonged to the last female Great Auk killed in 1844. Source: Cincinnati Museum
Researchers are collecting DNA samples for study: they drill a small hole in the base of the specimen and use a fine scraper to collect a sample from the foot, minimizing damage to the specimen.

Appearance #

  • The upper body and head are black, with large white patches in front of the eyes during summer. The eyes are very small, with hazel irises8.
  • It had a large, hard black beak with several grooves, and the inside of the beak was orange-red.
  • The upper neck was black like the head but slightly brownish, while the belly was white.
  • The wings were highly reduced, very small in size, with black feathers. The secondary feathers were white, and the side abdomen near the wings often appeared brown.
  • The upper tail coverts were black, and the lower tail coverts were white.
  • The feet were webbed, black, with small claws.
  • During the winter molt, the black of the front neck and chin gradually turned white, and the white patches around the eyes became lighter, eventually being replaced by black.

The Great Auk exhibited no sexual dimorphism. Some existing female specimens have a gray band on their sides, which some believe is unique to females9. Studies on Great Auk skeletons suggest that males might have been slightly larger, but this difference appears to be minimal10.

The black back feathers of the Great Auk were described as “like black satin”11.

The appearance of Great Auk chicks remains unknown12.

Left: Scientific illustration depicting the winter molt of the Great Auk and its egg. F. W. Frohawk et al. Public Domain
Right: Illustration of the Great Auk. Artist: Willem van der Merwe, from Deviantart
Rare winter Great Auk specimen.
Left: Natural History Museum of Copenhagen, killed in the winter of 1815 near Greenland. Photo by Brian Berg, sourced from (Kjartansdóttir 2019) (CC BY-NC-ND.4.0) | Right: Trinity College Museum, Dublin, Wikimedia Commons. Captured in May 1834 in the waters of Waterford Harbour, Ireland, it died after being kept in captivity for four months. This was the last sighting of a Great Auk in Ireland. For more information, see here and [10]. Notice the distinct molt marks on the neck.
Close-up of the winter Great Auk specimen at the Natural History Museum of Copenhagen, photographed by Peter Andrew Hosner, sourced from Twitter. The molt marks on the throat and "eye mask" are very clear.
Great Auk specimen, Museum of Natural History, Wrocław University (Muzeum Przyrodnicze we Wrocławiu), Poland
One of two Great Auk specimens preserved at this museum, possibly one of the very few specimens showing the Great Auk with spread wings
During the late stages of World War II, the museum was bombed during the Soviet siege of the Nazi German forces in Wrocław. The two Great Auk specimens were rescued and protected from the ruins
Source: Fundacja Collect (CC BY 4.0 DEED)
Great Auk juvenile specimen
Credit: Tyne and Wear Archives and Museums, part of the collection of the Natural History Society of Northumbria, sourced from Twitter
Currently housed at Great North Museum: Hancock, it is considered the youngest of the existing Great Auk specimens
Collecting DNA samples from a Great Auk specimen. Note the very small claws at the tips of the webbed feet.
Source: Cincinnati Museum

Life History #

Breeding and Rearing #

It is generally believed that, like similar auk species, the Great Auk was monogamous for life. Many accounts mention the Great Auk shaking its head vigorously, which was likely a courtship display13. During the breeding season, Great Auks were highly social, forming large and crowded colonies14. Because they were flightless, they had specific requirements for their breeding sites, needing gentle sloping rocks near the sea for easier access from the water. They were never observed landing on the mainland. However, the number of observed modern breeding sites was still significantly fewer than the theoretically suitable ones, for reasons that remain unclear. It might have been due to unknown food or location needs, or pressure to avoid human hunters15.

Great Auks did not build nests; they laid their eggs directly on the surface of the island rocks, such as open rock ledges, crevices, or at the base of coastal cliffs. Their mating sites and behaviors are unknown, but it is likely similar to their close relative, the Razorbill, which mates after coming ashore, with nesting sites not far from mating sites. Descriptions indicate that Great Auk pairs would “courageously” protect their eggs even in the face of being captured16.

Both sexes had brood patches, suggesting that parents likely took turns incubating. Recent studies found that Great Auks had two brood patches instead of the previously thought one, indicating they likely incubated eggs while lying flat rather than the previously hypothesized semi-upright posture17. Great Auk eggs were pear-shaped with markings similar to ink sketches. The shape likely helped them remain stable on sloped surfaces, preventing them from rolling off exposed rocks, and the markings helped parents identify their eggs in densely packed colonies.

Great Auks were vulnerable on land and thus minimized their time there. Their incubation period was about 39 to 44 days, with reports of as short as 26 days18. In comparison, the closely related Razorbill, with a much smaller body and egg, takes 35 to 37 days to incubate19. Likewise, the Great Auk’s rearing period was very short, with both parents feeding the chick on land. It is likely that within two weeks, or even 10 days, the chick followed its parents into the water and drifted in the North Atlantic, possibly traveling hundreds of kilometers from the shore before being fully fledged20.

When leaving the nest, Great Auk chicks weighed about 20% of an adult’s weight21. People have reported seeing Great Auk parents carrying their chicks into the sea on their backs22.

Great Auks during the breeding season, illustration
Artist: Heinrich Harder
Left: Cover of 'Great Auk Islands: A Field Biologist in the Arctic', depicting an imagined scene of Great Auk parents rearing a chick. The gray feathers on the chick likely come from (Otto Fabricius 1780)'s observations of the chick.
Right: Great Auks during the breeding season, illustration by John Gerrard Keulemans. This illustration contains some scientific inaccuracies: in the lower left center, three chicks surround one adult Great Auk, which is unlikely given the monogamous nature of the species. The size comparison between the chicks and the adult is also problematic, as the chicks would have gone to sea before reaching such a size. Furthermore, the chicks’ necks and cheeks are white, resembling the adult's winter molt, but with white eye patches, the basis why artist depicted them as such is unclear.
A great auk egg, illustration
Coloured lithograph by F. W. Frohawk, 1888. Wellcome Collection. Source: Wellcome Collection .
A great auk egg, Liverpool Museum collection. Source: Wikimedia Commons
This egg, covered in beautiful patterns, is considered the most beautiful among the 70+ Great Auk eggs preserved today, although its appearance is not very typical.
During the Victorian era, Great Auk eggs were highly prized collectibles. As the species became more endangered, wealthy individuals sought them even more eagerly, leading to an intensification of hunting to a much higher degree—a practice that seems utterly awful to today's people.

Diet #

The Great Auk primarily ate fish found in the shallow coastal waters of the North Atlantic, with a diet similar to today’s auk species in the region.

Naturalists who dissected Great Auks found their stomachs contained shorthorn sculpin (Myoxocephalus scorpius) and lumpsucker (Cyclopterus)23. The shorthorn sculpin often inhabits benthic zones, suggesting that the Great Auk likely had the ability to dive deep to catch fish.

Modern scientific analysis of Great Auk bones and soil from Funk Island identified the presence of the following North Atlantic fish, likely part of their diet24:

  • Atlantic menhaden
  • American shad
  • Capelin
  • Three-spined stickleback
  • North American bass (Morone), possibly White perch or small Striped bass
  • Flatfish

Studies suggest that Great Auk chicks might have fed on krill and other crustaceans, and that parents might have regurgitated food to feed their chicks, similar to many penguin species—another example of convergent evolution between these two unrelated species with similar appearances and vastly different fates25. Dissections of Great Auk chicks have found Rhodiola, a plant growing on arctic rocks23.

The last Great Auk in Ireland was kept alive in captive for four months26. It was found to prefer eating trout and other freshwater fish over sea fish. It also enjoyed mashed potatoes soaked in milk27.

A description of a live bird noted that a Great Auk could swallow a herring whole, and would eat three in succession to feel full11.

Great Auk illustration by John Gould, from "The Birds of Europe", 19th century, Public Domain

Distribution #

Confirmed modern breeding sites of the Great Auk include the following locations28:

  • Funk Island, off the eastern tip of Newfoundland, Canada, Labrador Sea

This island was originally known as “Penguin,” this name was first given to the Great Auk and later to penguins29.

Funk Island was the largest modern breeding site of the Great Auk. When Europeans arrived, tens of thousands of Great Auks were breeding there during the season, with estimates exceeding 100,000 pairs30. By the beginning of the 19th century, Great Auks on Funk Island were nearly extinct.

In 1887, Frederick A. Lucas, then curator of the National Museum of Natural History in the United States, visited Funk Island. The expedition team found a large number of Great Auk remains in the island’s soil and excavated some of them. Nearly all museum collections of complete Great Auk skeletons come from Funk Island. Remains of iron pots, likely used to boil Great Auk bodies for easier feather removal, were also found in the upper soil layer. Frederick estimated that millions of Great Auks perished on Funk Island. The expedition also discovered that many Atlantic puffins (Fratercula arctica) nested on Funk Island, which had not been seen before the Great Auk’s extinction. It is speculated that the discarded bodies of Great Auks created a nutrient-rich soil suitable for puffin burrows31.

Scientists also found stone enclosures near the Great Auk breeding grounds on Funk Island, which were likely used to corral and slaughter the birds32.

One of Newfoundland’s early pioneers, British explorer George Cartwright(1739 - 1819), recorded the exploitation of Great Auks on Funk Island:

Innumerable flocks of sea fowl breed upon it every summer which are of great service to the poor inhabitants of Fogo; who make voyages there to load with birds and eggs… But it has been customary of late years for several crews of men to live all the summer on that island, for the sole purpose of killing birds for the sake of their feathers; the destruction which they have made is incredible. If a stop is not soon put to that practice the whole breed will be diminished to almost nothing, particularly the penguins: for this is now the only island they have left to breed upon. 33

  • Papa Westray Island, part of the Orkney Islands, northeastern Scotland

In 1813, a male Great Auk was killed on Papa Westray Island in the Orkney Islands. For several years prior, this Great Auk had been attempting to breed in the area with a female. They were known to locals as the “King and Queen of the Auks,”34.

Other islands in the Orkney archipelago were thought to be suitable for Great Auk breeding, but no confirmed sightings were recorded. It is speculated that the two Great Auks seen on Papa Westray were strays from other breeding colonies35.

  • St Kilda, an archipelago in the North Atlantic west of Scotland

In July 1840, five fishermen from St Kilda captured a Great Auk sleeping on a rock ledge on Stac an Armin. They tied its feet with a rope and kept it in captivity. Three days later, during a storm, the frightened fishermen believed it might be a witch and stoned the Great Auk to death36.

This was the last Great Auk recorded in the British Isles. Recent studies suggest this event might have occurred in 1848, rather than the previously thought 1840. If true, this would make it the last Great Auk in the world, instead of the pair killed in 1844 on Eldey Island37.

Although there are early documented records of Great Auk breeding colonies on St Kilda, many aspects remain unclear because the island’s coastal areas are steep and seem unsuitable for Great Auk landings38.

  • Geirfuglasker and Eldey Island, off the southwestern coast of Iceland

Geirfuglasker, near the Reykjanes Peninsula in Iceland, was one of the last refuges of the Great Auk. The island was surrounded by turbulent waters, making human access difficult. In 1813, during the War of the Sixth Coalition, when Denmark was allied with France and opposed to Britain, a Danish ship searching food around Iceland landed on Geirfuglasker and killed a large number of Great Auks39.

In 1830, an underwater volcanic eruption and subsequent earthquakes caused Geirfuglasker to sink below sea level, forcing the Great Auks to move to Eldey Island, which was more accessible to humans.

On June 3, 1844, two Great Auks incubating an egg were killed on Eldey Island, and the egg was broken. This is generally considered the last two Great Auks on Earth.

Other islands around Iceland also had sporadic records of Great Auk breeding, with Grímsey Island in the north being notable40.

  • Bird Rock (Rochers aux Oiseaux), Magdalen Islands, Quebec, Canada, in the Gulf of St. Lawrence, near Newfoundland

Great Auk breeding colonies were observed here from the 16th to the 17th centuries. There were still sizeable colonies in the late 17th century, but by 1700, this breeding colony had disappeared, with most individuals likely migrating to Funk Island41.

It is speculated that the Great Auk population here migrated from Penguin Island near Cape La Hune on the southern side of Newfoundland. This breeding site seems to have been abandoned earlier, but the name indicates it was once a breeding ground for Great Auks42.

Other potential North American breeding sites for Great Auks include Cape Cod in Massachusetts and areas near Cape Breton in Nova Scotia, Canada. They were also sighted in Labrador43.

  • (?) Faroe Islands, an archipelago in the North Atlantic, north of the British Isles and east of Iceland, currently an autonomous territory of Denmark

There have been sporadic records of Great Auk sightings and captures in the Faroe Islands, the last being in July 180844. However, there was never a large breeding colony of Great Auks here, nor were there confirmed egg-laying sites. It is speculated that the Great Auks appearing in the Faroe Islands were immature birds arriving from North America via ocean currents. These birds likely escaped the massacres at Funk Island but instinctively attempted to return to breed, using the Faroe Islands as a stopover45.

The term “Northern Penguin” for the Great Auk is based on early European naturalists’ misconceptions, incorrectly suggesting that the Great Auk inhabited the Arctic. There is no reliable evidence that the Great Auk’s modern range included the Arctic Circle. In 1821, there was a report of a Great Auk being captured on Disko Island in Greenland, which is the northernmost known occurrence of the Great Auk. However, the reliability of this record is highly questionable46. Early Eskimo settlements in Qeqertasussuk, Greenland, have also unearthed Great Auk fossils47. While the Great Auk’s presence in the southern edge of the Arctic Circle is undeniable, it is difficult to say they had a stable habitat and breeding ground there.

The Great Auk’s prehistoric range was broader than the modern records indicate, extending to lower latitudes. In North America, 14th-century Great Auk bones have been found in Florida48. In Southern Europe and North Africa, Holocene Great Auk fossils have been unearthed in Iberia, the Gibraltar region, and countries like Spain, Italy, France, and Morocco, dating back several thousand years BC. These fossils often show evidence of human hunting and butchering, indicating that early humans were already hunting Great Auks49.

Further north, there are numerous records of Great Auk fossils found outside their modern range, such as in Norway, Sweden, and Denmark on the Scandinavian Peninsula. The Norwegian population possibly survived until after AD 1000, with their range extending east to Finnmark bordering Russia50. The Dutch coast also had Great Auk populations as late as the Roman Empire51.

It can be concluded that the Great Auk was once widely distributed across the Atlantic Ocean. Their migration patterns are unknown. Generally, they returned to the sea after breeding in the summer, following ocean currents to warmer regions in winter. Whether they had a migratory rhythm like other birds is unknown, as they were flightless and likely highly dependent on ocean currents52. Their populations began declining before recorded history due to climate change and human hunting. However, the impact of human hunting became more significant over time53, with hunting pressure far exceeding the effects of climate change. Evidence suggests that Great Auk populations suffered a dramatic decline after the advent of the Iron Age54.

Distribution range and breeding sites of the Great Auk. Blue represents historical distribution range, yellow dots indicate breeding sites.
Breeding sites from left to right: Bird Rock in the Magdalen Islands; Penguin Island near Cape La Hune; Funk Island; southern Greenland; ?; Geirfuglasker and Eldey Island; St Kilda; Faroe Islands; Papa Westray Island
There are some uncertainties in this map. The mark near the upper part of Iceland is unclear, although there were likely other breeding sites around Iceland besides Geirfuglasker and Eldey Island. Also, while there are records of sightings and captures in southern Greenland, it is hard to say that the Great Auk regularly bred there. See (Gaskell 2000), p148 for a discussion.
Map by Shyamal. Wiki Commons , Public Domain
In a study by (Jessica Emma Thomas 2019), researchers used two GPS drifting capsules to simulate possible migration routes of Great Auks along ocean currents in the North Atlantic.
Red stars: Great Auk breeding sites; green and yellow lines: routes of the two GPS drifting capsules; red arrow lines: warm ocean currents (Gulf Stream and North Atlantic Current); blue arrows: cold ocean currents (East Greenland Current and Labrador Current)
Source: Researchgate (CC BY 4.0)
A Last Stand by Errol Fuller. Source: Wikimedia Commons
Left: Extinct Animals series, drawn by Zhou Sheji, Zcool
Right: Oil painting by John Duncan (c.1846–after 1907), The Natural History Society of Northumbria Great North Museum: Hancock, ArtUK, used non-commercially under usage terms

Other #

There are no direct records of the natural lifespan and reproductive age of the Great Auk. Based on the Razorbill, it is estimated that their lifespan was about 20-25 years, with first breeding occurring at 4-7 years of age13.

Like similar birds such as the Razorbill, and based on limited observations of the Great Auk, their long, hard beaks were likely a secondary sexual characteristic used for courtship displays and a primary tool for communication within the group. There are records indicating that captive Great Auks would keep their beaks open, signaling a readiness to attack55. The powerful beaks of Great Auks may have enabled them to easily crush the shells of their prey. One account from a hunt on Geirfuglasker mentions a desperate Great Auk biting a man’s forearm through a leather jacket, causing blood to flow down his sleeve56.

In normal circumstances, the Great Auk’s call was a low murmur. Records show that captive Great Auks would raise their heads, shake their heads and necks, and make this sound to express a desire for food when being fed by humans57. However, when threatened, the Great Auk could produce a very loud call, reportedly louder than the famously noisy Northern Gannet55.

Great Auks were exceptionally skilled swimmers. Accounts describe them immediately diving underwater when frightened, rather than fleeing along the surface58. In addition to their webbed feet, they used their reduced, flightless wings for propulsion and balance underwater, functioning similarly to fins59. When capturing the last British specimen near Papa Westray Island, it was documented that it took several hours of chasing in a six-oared boat before before people gave up capturing or shooting it in the water60.

Observers believed that Great Auks had poor vision on land but excellent hearing, being very sensitive to noise. It was also speculated that the misconception about their poor land vision might be due to people not recognizing those birds’ nictitating membranes61.

Confirmed diving depths for Great Auk hunting ranged from 50 to 100 meters. Based on their size and comparisons to similar species, they may have been capable of diving even deeper, possibly up to nearly 200 meters62.

19th-century illustrations of the Great Auk
Left: Die Gartenlaube. Ernst Keil's Nachfolger. 1884. Public Domain.
Right: Lost and vanishing birds;. London, J. Macqueen, 1898. CC BY 2.0 DEED.
Illustration of Great Auks. The two Great Auks in the background seem to be engaged in a courtship struggle, with one threatening the other with its beak. This is quite interesting and scientifically accurate.
Popular Science Monthly Volume 62. The great auk. between 1902 and 1903. Unknown author. Public Domain.

Accounts #

  • In 1534, French explorer Jacques Cartier was on his first voyage to North America. He was the first European to travel inland in North America, although his original mission from the King of France was to find a passage to the East. He observed Great Auks on “Bird Rock” in the Magdalen Islands of Quebec. This is one of the earliest records of the Great Auk.

    And the twenty-first day of the said month of May we departed from the said harbor with a west wind, and were borne to the north a quarter northeast from Cape Bonavista as far as to the Isle of Birds, the which island was all sur- rounded and inclosed by a bank of ice, rent and shattered in pieces. Notwithstanding the said bank, our two boats went to the said island in order to get some of the birds, of which there is so great a number that it is an incredible thing to one who does not see it; for, notwithstanding that the said island comprises about a league in circumference, it may be so very full of them that it seems as if one had stowed them there. There are a hundred times more of them round about it and in the air than within the island, of which a part of these birds are large as geese, black and white, and have a beak like a crow, and are always in the sea, without ever being able to fly in the air, forasmuch as they have little wings like the half of one, wherewith they fly as stoutly in the sea as the other birds do in the air, and these birds are so fat that it is a marvelous thing. We named these birds Apponatz63 with which our two boats were loaded in less than half an hour, as it were with stones, wherefore each of our ships salted four or five butts 64 of them, besides what we could eat of them fresh.

  • In 1583, explorer Edward Haies joined Humphrey Gilbert’s fleet to the Newfoundland Islands and observed numerous Great Auks on Funk Island.

    Forsaking this bay and uncomfortable coast (nothing appearing unto us but hideous rocks and mountains, bare of trees, and void of any green herb) we followed the coast to the south, with weather fair and clear. We had sight of an island named Penguin29, of a fowl there breeding in abundance almost incredible, which cannot fly, their wings not able to carry their body, being very large (not much less than a goose) and exceeding fat, which the Frenchmen use to take without difficulty upon that island, and to barrel them up with salt. But for lingering of time, we had made us there the like provision. 65

  • In the late 16th century, English colonist Richard Whitbourne was involved in the fishing industry near Newfoundland. He later described the Great Auk as follows:

    These Penguins are as bigge as Geese, and flye not, for they haue but a little short wing, and they multiply so infinitly, vpon a certaine flat Iland, that men driue them from thence vpon a boord, into their boates by hundreds at a time; as if God had made the innocency of so poore a creature, to be∣come such an admirable instrument for the susten∣tation of man.66

  • In 1697, Scottish writer Martin Martin visited the isolated St Kilda archipelago in the North Atlantic and encountered Great Auks there:

    The Sea-Fowls are, first, Gairfowl, being the stateliest, as well as the Largest of all the Fowls here, and above the Size of a Solan Goose, of a Black Colour, Red about the Eyes, a large White Spot under each Eye, a long broad bill; stands stately, its& whole body erected, its Wings short, it flyeth not at all, lays its egg upon the bare Rock, which, if taken away, it lays no more for that year; it is Palmipes67, or Whole-Footed, and has the Hatching spot upon its Breast, i.e. a bare Spot from which the Feathers have fallen off with the heat in Hatching; its egg is twice as big as that of a Solan Goose, and is variously spotted, Black, Green, and Dark; it comes without regard to any Wind, appears the first of May68, and goes away about the middle of June.69

  • In the 17th century, French-born merchant and North American colonist Nicolas Denys observed Great Auks on the eastern coast of North America:

    The Great Auk (Pennegoin) is another bird, variegated in white and black. It does not fly. It has only two stumps of wings with which it beats upon the water to aid in fleeing or diving. It is claimed that it dives even to the bottom to seek its prey upon the Bank. It is found more than a hundred leagues 70 from land, where, nevertheless, it comes to lay its eggs, like the others. When they have had their young, they plunge into the water; and their young place themselves upon their backs, and are carried like this as far as the Bank. There one sees some no larger than chickens, although they grow as large as geese. All those birds are good to eat by the fishermen. As for myself I do not find them agreeable. They taste of oil because of the quantity of fish and of livers they eat; and they serve to make fish oil. The fishermen collect them for this purpose. There are vessels which have made as much as ten or twelve puncheons 71 of it. This is nearly everything which is practiced in the fishery for green Cod upon the Grand Bank. 22

  • From 1794 to 1795, Royal Navy seaman Aaron Thomas (1762 - 1799) served aboard HMS Boston, reaching Newfoundland, where he participated in the mass slaughter of Great Auks on Funk Island. He recorded his experiences as follows72:

    Funk Island is a barren spot inhabited only by Penguins and other Birds. The astonishing quantity which resort to this Island is beyond beliefe of any person, except those who have witnessed something similar in other parts. As soon as you put your foot on shore you meet with such Thousands of them that you cannot find a place for your feet and they are so lazy that they will not attempt to move out of your way. If you come for their Feather you do not give yourself the trouble of killing them, but lay hold of one and pluck off the best of the Feathers. You then turn the poor Penguin adrift, with his Skin half naked and torn off, to perish at his leasure. This is not a very humane method, but it is the common Practize.

    If you go to the Funks for Eggs, to be certain of getting them fresh, you pursue the following Rule: - you drive, knock and shove the poor Penguins in Heaps! You then scrape all the Eggs in Tumps in the same manner you would a Heap of Apples in an Orchard in Herefordshire. Numbers of these Eggs, from being dropped some time, are stale and useless, but you having cleared a space of ground the circumference of which is equal to the quantity of Eggs you want, you retire for a day or two behind some Rock at the end of which time you will find plenty of Eggs - fresh for certain! - on the place where you had before cleared.73

    While you abide on this island you are in the constant practize of horrid cruelties for you not only Skin them Alive, but you burn them Alive also to cook their Bodies with. You take a kettle with you into which you put a Penguin or two, you kindle a fire under it, and this fire is absolutely made of the unfortunate Penguins themselves. Their bodys being oily soon produce a Flame; there is no wood on the island.74

    I had the following information from a person in St. John’s… ‘About twenty years ago when this kind of Traffick was Lawfull, I made two Trips to the Funks. In these Trips I gathered, with one person with me, half a Ton of Feathers and as many Eggs as sold in St. John’s for Thirty Pounds!’

    This skinning and taking the Eggs from the Funks is now prohibited and they are allowed to take the Birds only for Bait to catch Fish with. [But] about three years ago some fellows were detected in this kind of Plunder. They were brought to St. John’s and flogged at a Cart’s Tail. But I am told there is a quantity of Feathers [still] purloined from these Islands every year.”

  • In 1821, Scottish naturalist John Fleming observed a Great Auk captured near the St Kilda archipelago. This might be the only professional record of a live Great Auk:

    When on the eve of our departure from this island, we got on board a live example of the Great Auk (Alca impennis) which Mr Maclellan, the tacksman [i.e. tenant] of Glass had captured some time before off St Kilda. It was emaciated. and had the appearance of being sickly, but, in the course of a few days, it became sprightly, having been plentifully supplied with fresh fish, and permitted occasionally to sport in the water with a cord fastened to one of its legs to prevent escape. Even in this state of restraint it performed the motions of diving and swimming under water with a rapidity that set all pursuit from a boat at defiance. A few white feathers were at this time making their appearance on the sides of its neck and throat, which increased considerably during the following week and left no room to doubt that, like its congeners, the blackness of the throat feathers in summer is exchanged for white during the winter season. I may add that the black colour of the throat of the Razor-bill (Alca torda) was at this time undergoing a similar change.75

  • In 1858, British naturalist John Wolley (1823 - 1859) and Alfred Newton (1829 - 1907) searched for traces of the Great Auk near Iceland and interviewed the three Icelandic villagers who killed the last pair of Great Auks on Eldey Island on June 3, 184476. John Wolley died young the year after this visit, and the findings were later published by Alfred Newton, including records of the circumstances surrounding the death of the last two Great Auks.

    Tlie Garc-fowls sliowed not the slightest disposition to repel tlic invadws, but itnniediately ran along under the high cliff, their lrcads ervct, their little wings sotnewhat extended. They uttered no cry of alarm, and moved, with their short steps, about as quickly as a man could walk. Jón with outstretched arms drove one into a corner, where he soon had it fast. Sigurður and Ketill pursued the second, and the former seized it close to the edge of the rock, here risen to a precipice some fathoms high, the water being directly below it. Ketill then returned to the sloping shelf whence tlie birds had started, and saw an egg lying on the lava slab, which he knew to be a Garc-fowl’s. He took it up, but finding it was broken, put it down again. Whether there was not also another egg is uncertain, All this took place in much less time than it takes to tell it. They hurried down again, for the wind was rising. The birds were strangled and cast into the boat, and the two younger man followed.77

    Wolley’s notes include his interviews with the three villagers: Sigurðr Islefsson, Jón Brandsson, and Ketil Ketilsson78. The two Great Auks tried to escape separately, and Sigurðr Islefsson and Ketil Ketilsson each strangled one. Sigurðr Islefsson’s recollection is as follows:

    The rocks were covered with blackbirds 79 and there were the Geirfugles … They walked slowly. Jón Brandsson crept up with his arms open. The bird that Jón got went into a corner but [mine] was going to the edge of the cliff. [I] caught it close to the edge - a precipice many fathoms 80 deep. The black birds were flying off. I took him 81 by the neck and he flapped his wings. He made no cry. I strangled him. 82

    My bird ran [from where it was first seen] about 20 fathoms. It walked like a man … but moved its feet quickly. It began to move as the people came on the rock and as the blackbirds began to fly. Its wings lay close to its sides - not hanging out. As it was held by the neck it hung its wings out a little. 82

    Ketil Ketilsson’s recollection is essentially the same:

    [We] ran together after the birds, but as [we got near the edge of the precipice my] head failed me and I stopped; Sigurd went on and seized the bird … This one must have run quicker than a man can walk . It held itself quite straight up, with its wings close to its sides, and as it ran it made no noise or cry. 82

  • In 1883, Norwegian fisherman Herr Brodtkorb told Oslo University researcher Robert Collet about his sighting of four Great Auks at sea near Vardö, Finnmark County, in April 1848. He claimed to have shot one of them. This sighting is not widely accepted by the scientific community due to a lack of direct evidence, as Herr Brodtkorb claimed that the bird’s body was washed away by the tide. Additionally, his description suggests it could have been a misidentification of the common loon (Gavia immer), and Vardö is at a much higher latitude than the known modern range of the Great Auk83. Herr Brodtkorb described the Great Auks as follows:

    In swimming they used both wings and feet, and also dived, but did not stay long under the water. It almost seemed as if they only went through the tops of the waves. The birds kept together and did not seem afraid. We .. heard a cry… they emitted when they drew more closely together. It resembled a cackling, as if they wished to call one another … When the shot went off all the four birds disappeared; but shortly after I saw the remaining three paddling on farther until they disappeared behind the surging waves.84

Skeleton of Great Auk, excavated in Newfoundland, May 1958. Source:Florida Museum, Photo by Kristen Grace
Skeleton of Great Auk
Left: Photo by Rhett A. Butler. butlernature.com
Right: Smithsonian Institution Archives, Acc. 11-007, Box 018, Image No. MNH-4284. Smithsonian Institution Archives. Public Domain
Specimen of Great Auk, Natural History Museum of Oslo, Norway
Credit: Karsten Sund/Natural History Museum of Oslo. Endangered nature – threats against biodiversity
Great Auk
Artist: Ida Kalsta

  1. Moum T, Arnason U, Árnason E. 2002. Mitochondrial DNA Sequence Evolution and Phylogeny of the Atlantic Alcidae, Including the Extinct Great Auk (Pinguinus impennis). Molecular Biology and Evolution. 19(9):1434–1439. doi:https://doi.org/10.1093/oxfordjournals.molbev.a004206. The Razorbill is the only extant species of the genus, but recent fossil research has discovered more species from the Pliocene. See: Smith NA, Clarke JA. 2011. An Alphataxonomic Revision of Extinct and Extant Razorbills (Aves, Alcidae): A Combined Morphometric and Phylogenetic Approach. Ornithological Monographs. 72(1):1–61. doi:https://doi.org/10.1525/om.2011.72.1.1. Pliocene fossils also revealed another species of the genus Pinguinus, distributed on the western Atlantic coast, which went extinct in ancient times, named Pinguinus alfrednewtoni. It is generally believed that they and the Great Auk were sister species that diverged due to geographic isolation. After Pinguinus alfrednewtoni’s extinction, the Great Auk, originally distributed on the eastern Atlantic coast, migrated across the sea to occupy similar ecological niches on the west coast. See: Ray CE, Bohaska DJ. 2001. Geology and Paleontology of the Lee Creek Mine, North Carolina, III. p279. ↩︎

  2. Another ancient relative of the Great Auk mentioned in [1] was slightly larger than the Great Auk. ↩︎

  3. Most specimens and illustrations are based on the Great Auk in summer, depicting their oval “eye patches” (actually in front of the eyes). This is because Great Auks lived at sea in winter and only came ashore to breed in summer, giving humans the opportunity to encounter (and hunt them to extinction). Winter specimens are rare. In the winter of 1815, a Great Auk was killed in the sea off Fiskenaesset (now Qeqertarsuatsiaat) in Danish Greenland, and its taxidermied specimen is preserved in the Natural History Museum of Copenhagen. The illustrations on this page reference: Kjartansdóttir K. 2019. The changing symbolic meaning of the extinct great auk and its afterlife as a museum object at the Natural History Museum of Denmark. Nordisk Museologi. 26(2):41–56. doi:https://doi.org/10.5617/nm.7476. Photographs of the same specimen by Geert Brovad can also be found in Meldgaard M. 1988. The Great Auk, Pinguinus impennis (L.) in Greenland. Historical Biology. 1(2):145–178. doi:https://doi.org/10.1080/08912968809386472. and Morten Meldgaard. 2004. Ancient harp seal hunters of Disko Bay (Vol. 330): Subsistence and settlement at the Saqqaq culture site Qeqertasussuk (2400-1400 BC), West Greenland. Museum Tusculanum Press. ↩︎

  4. The skins and internal organs of the two Great Auks were sent to Denmark, but the skins were later lost. Scientists confirmed through DNA analysis of the organs that the skin of the last male Great Auk is preserved in a specimen at the Royal Belgian Institute of Natural Sciences. See: Thomas JE, Carvalho GR, Haile J, Martin MD, Samaniego Castruita JA, Niemann J, Sinding M-HS, Sandoval-Velasco M, Rawlence NJ, Fuller E, et al. 2017. An ‘Aukward’ Tale: A Genetic Approach to Discover the Whereabouts of the Last Great Auks. Genes. 8(6). doi:https://doi.org/10.3390/genes8060164. The whereabouts of the female skin remain unknown. Researchers speculate that the skin of a Great Auk purchased by the Cincinnati Museum from a London dealer in the 20th century may belong to the female killed in 1844. The Cincinnati Museum stated in a 2020 publication that the same research team requested Great Auk tissue for DNA sequencing, and they complied. As of 2024, I have not found the results of this sequencing. See here↩︎

  5. The bounty was offered by Icelandic merchant Carl Franz Siemsen, an agent for several museums. ↩︎

  6. Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p144. ↩︎

  7. From ornithologist Henry Drummond-Hay. The IUCN Red List includes this sighting, seemingly based mainly on the credibility of the observer. For critiques of this sighting, see this blog post↩︎

  8. Based on observations of live birds. See Fuller Errol. The Great Auk. 1999. p25. ↩︎

  9. Walter Rothschild. Extinct Birds. London: Hutchinson & Co., 1907. ↩︎

  10. Livezey, Bradley C. “Morphometrics of flightlessness in the Alcidae.” The Auk 105.4 (1988): 681-698. ↩︎

  11. Ole Worm - Olaus Wormius. Museum Wormianum. 1655. Referenced in Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p65. ↩︎ ↩︎

  12. A juvenile specimen preserved at the Great North Museum: Hancock in Newcastle is considered the youngest Great Auk specimen, although its plumage characteristics seem similar to adults. See: https://www.theguardian.com/environment/2014/dec/10/country-diary-susie-white-museum-newcastle-great-auk. A specimen preserved at the Národní Muzeum in Prague is also considered juvenile. See (Fuller, 1999). ↩︎

  13. Bengtson, Sven-Axel. “Breeding ecology and extinction of the Great Auk (Pinguinus impennis): Anecdotal evidence and conjectures.” The Auk 101.1 (1984): 1-12. ↩︎ ↩︎

  14. A recollection mentioned that Great Auks bred “close under the rocks and sitting so that they were shoulder to shoulder.” See Fuller Errol. The Great Auk. 1999. p39. ↩︎

  15. Human factors were likely the most important, as all modern nesting sites of the Great Auk were far from “civilized” areas. See the discussion in (Fuller, 1999). ↩︎

  16. During the 1829 killing of two Great Auks on Geirfuglasker Island in Iceland. (Nils Kjærbölling, 1856), referenced in Fuller, 1999, p136. However, according to p218, this account is not very certain. ↩︎

  17. Birkhead T R, Fiebig J, Montgomerie R, et al. The Great Auk (Pinguinus impennis) had two brood patches, not one: confirmation and implications[J]. Ibis, 2022, 164(2): 494-504. The hypothesis of semi-upright incubation can be found in Birkhead T, Russell D, Garbout A, et al. New insights from old eggs–the shape and thickness of Great Auk Pinguinus impennis eggs[J]. Ibis, 2020, 162(4): 1345-1354. ↩︎

  18. Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p153. ↩︎

  19. The average length of Razorbill eggs is about 7.3 cm, while Great Auk eggs exceed 12 cm. ↩︎

  20. Fisher J, Lockley R M. Sea-birds: An Introduction to the Natural History of the Sea-birds of the North Atlantic[J]. 1954. ↩︎

  21. Houston A I, Wood J, Wilkinson M. How did the Great Auk raise its young?[J]. Journal of evolutionary biology, 2010, 23(9): 1899-1906. ↩︎

  22. Nicolas Denys. The Description and Natural History of the Coasts of North America. 1672. Cited from Allen, E. G. (1939). Nicolas Denys, a Forgotten Observer of Birds. The Auk, 56(3), 283–290. doi:10.2307/4079049 ↩︎ ↩︎

  23. Otto Fabricius’ study on Great Auks around Greenland. Fabricius, Otto. Favna groenlandica. (1780). Cited from Fuller Errol. The Great Auk. 1999. p44. ↩︎ ↩︎

  24. Olson SL, Swift CC, Mokhiber C. 1979. An Attempt to Determine the Prey of the Great Auk (Pinguinus impennis). The Auk. 96(4):790–792. ↩︎

  25. Hobson KA, Montevecchi WA. 1991. Stable isotopic determinations of trophic relationships of great auks. Oecologia. 87(4):528–531. doi:https://doi.org/10.1007/bf00320416. ↩︎

  26. Perhaps one of the very few Great Auks to die of natural causes after being discovered by humans, although it did not live long after being captured. ↩︎

  27. Thomas Parkin. The Great Auk, Or Garefowl (Alca Impennis, Linn.). 1911. Also see: Fuller Errol. The Great Auk. 1999. p57. ↩︎

  28. Fisher J, Lockley R M. Sea-birds: An Introduction to the Natural History of the Sea-birds of the North Atlantic[J]. 1954. p66. See another earlier list by Rothschild, which is basically the same but additionally lists the Faroe Islands: Rothschild, Walter. Extinct Birds. 1907. Recent research can be found in: Søren Bay Kruse Thomsen. The History of the Decline and Fall of the Great Auk. 2023. https://www.theextinctions.com/articles-1/the-history-of-the-decline-and-fall-of-the-great-auk ↩︎

  29. The name “Penguin” was later borrowed for the penguin species. The etymology of the word is not entirely clear. One theory suggests it comes from the Welsh word pen gwyn for “white head,” referring to the Great Auk’s white eye patches in summer; another theory is that it is similar to the Latin word pinguis for “fat,” referring to the Great Auk’s body shape. ↩︎ ↩︎

  30. Montevecchi, William A., and Leslie M. Tuck. “Newfoundland birds: exploitation, study, conservation.” (1987). p147. ↩︎

  31. Lucas F A. The expedition to Funk Island, with observations upon the history and anatomy of the Great Auk[J]. Report of the United States National Museum for the year ending June 30, 1888 (Pt. 2 of the Annual Report of the Board of Regents of the Smitsonian Institution for the year ending June 30, 1888.), 1888. ↩︎

  32. (Peter Stuwitz 1841), Cited from: funkisland.ca: https://funkisland.ca/scientific-history/ ↩︎

  33. Recorded on July 25, 1785, when a ship loaded with Great Auks sailed from Funk Island to St. John’s. George Cartwright witnessed this scene. Cited from (Gaskell 2000), the author suggests that the sailors’ indiscriminate killing might be related to material shortages during the American Revolutionary War. ↩︎

  34. The male Great Auk specimen was preserved and is now housed in the Natural History Museum in Tring, UK. It is one of the few confirmed male specimens collected in the British Isles. For the specimen’s provenance, see (Fuller, 1999). It is believed that Papa Westray Island itself was unsuitable for Great Auk egg-laying, and it is speculated that they bred on a nearby islet. Archaeological records indicate that the Great Auk had a wider historical distribution in the British Isles. See: Yalden, Derek, and Umberto Albarella. The history of British birds. Oxford University Press, 2009. ↩︎

  35. Fuller Errol. The Great Auk. 1999. p362. Holm Island is considered a possible breeding site. ↩︎

  36. This event has multiple versions, mainly from the oral recollections of one of the fishermen, Lachlan McKinnon. See: Fleming A. The last of the great auks: oral history and ritual killings at St Kilda[J]. Scottish Studies, 2024, 40: 29-40. ↩︎

  37. John A. Love, ‘The Last Great Auk?’ in Scottish Birds 31, no. 4 (2011), 346; See also: Historical Rare Birds. https://www.historicalrarebirds.info/cat-ac/great-auk; GAREFOWL: https://garefowl.co.uk/great-auk ↩︎

  38. Fuller Errol. The Great Auk. 1999. p366. ↩︎

  39. (Newton, Alfred, 1861). Cited from Gísli Pálsson. 2024. The Last of Its Kind. Princeton University Press. ↩︎

  40. (Gaskell, 2000) compiled the historical breeding sites of Great Auks in Iceland. See: Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p18. ↩︎

  41. Attention frag’îles (Association). Les oiseaux des Îles-de-la-Madeleine: populations et sites d’observation[M]. L’Étang-du-Nord, Québec: Attention frag’îles, 1992. See also: Les Oiseaux du Québec. https://www.oiseauxqc.org/details.jsp?no=1750 and Cokinos C. 2009. Hope Is the Thing With Feathers. J P Tarcher. p319. ↩︎

  42. Lucas, Frederic A. “Great Auk Notes.” The Auk 5, no. 3 (1888): 278-283. See also: Raghnall MacilleDhuibh. Why are penguins so called. 2006. https://www.querndust.co.uk/PDFs/530Penguin2.pdf ↩︎

  43. Grieve, S. “The Great Auk, or Garefowl. Its History, Archaeology and Remains, Digitally Printed Version 2015 ed.” (1885). Also see (Thomsen 2023) and (Gaskell 2000). ↩︎

  44. H.W. Feilden. Birds of the Faeroe Islands. 1872. Cited from (Gaskell 2000). ↩︎

  45. Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p119. Although the migration routes of the Great Auk are uncertain, recent DNA sequencing has proven that the Great Auks on both the east and west coasts of the Atlantic were a closely connected population. See: Thomas, Jessica E., Gary R. Carvalho, James Haile, Nicolas J. Rawlence, Michael D. Martin, Simon YW Ho, Arnór Þ. Sigfússon et al. “Demographic reconstruction from ancient DNA supports rapid extinction of the great auk.” Elife 8 (2019): e47509. ↩︎

  46. (Benicken 1824). Cited from (Gaskell 2000). Ironically, this understanding may have lessened the psychological burden on humans when they slaughtered Great Auks in the 18th and 19th centuries—they subconsciously believed that more Great Auks existed in the far north. ↩︎

  47. (Morten Meldgaard 2004). See[3]. ↩︎

  48. Pierce Brodkorb, Great Auk and Common Murre from a Florida Midden, The Auk, Volume 77, Issue 3, 1 July 1960, Pages 342–343, https://doi.org/10.2307/4082490 ↩︎

  49. Campmas E, Laroulandie V, Michel P, et al. 22 A great auk (Pinguinus impennis) in North Africa: d iscovery of[C]//Birds in Archaeology: Proceedings of the 6th Meeting of the ICAZ Bird Working Group in Groningen (23.8-27.8. 2008). Barkhuis, 2010, 12: 233. There is substantial evidence of early human hunting of Great Auks. Over 200 Great Auk beaks were found in a Native American grave in Newfoundland. See: Tuck J A. Ancient people of Port au Choix: the excavation of an Archaic Indian cemetery in Newfoundland[J]. (No Title), 1976. ↩︎

  50. Hufthammer A K, Hufthammer K O. The great auk in Norway: From common to locally extinct[J]. International Journal of Osteoarchaeology, 2023, 33(4): 588-597. The 1848 sighting of four Great Auks at sea also occurred in this area, possibly the last remnants of the population. ↩︎

  51. Groot M. The great auk (Pinguinus impennis) in the Netherlands during the Roman period[J]. International Journal of Osteoarchaeology, 2005, 15(1): 15-22. See also: Langeveld B W. New finds, sites and radiocarbon dates of skeletal remains of the Great Auk Pinguinus impennis from The Netherlands[J]. Ardea, 2020, 108(1): 5-19. ↩︎

  52. Refer to (Langeveld 2020) for an examination of the Great Auk’s wintering on the Dutch coast. ↩︎

  53. This does not imply that early indigenous hunting practices were moderate in intention, aiming to preserve Great Auk populations. For a similar viewpoint, see: Cokinos C. 2009. “Hope Is the Thing With Feathers.” J P Tarcher. p317. This is a myth not limited to the Great Auk; it has appeared in the context of many extinct animals, but may not reflect reality. The modern habitats of the Great Auk were very remote islands, which suggests they did not voluntarily restrict themselves to such isolated locations. A burial site of indigenous people in Newfoundland contained over 200 Great Auk beaks, which hardly suggests sustainable exploitation. ↩︎

  54. Refer to (Thomsen 2023) for a discussion. ↩︎

  55. Fuller Errol. The Great Auk. 1999. p56. ↩︎ ↩︎

  56. Benicken, J. C. “Beyträge zur nordischen Ornithologie.” Isis von Oken 2.8 (1824): 877-891. Cited from (Gaskell 2000). ↩︎

  57. John Fleming’s observation of a living bird. See: Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p13. ↩︎

  58. Newton, Alfred. “XLII.—Abstract of Mr. J. Wolley’sResearches in Iceland respecting the Gare‐fowl or Great Auk (Alea impennis, Linn.).” Ibis 3, no. 4 (1861): 374-399. ↩︎

  59. Refer to Nicolas Denys’ accounts. The analogy of fins comes from Colonel Montagu, cited in (Gaskell 2000). ↩︎

  60. William MacGillivray. A History of British Birds, indigenous and migratory. 1852. Cited from (Fuller, 1999). Also see: BritishBirds. 2022. https://britishbirds.co.uk/content/focus ↩︎

  61. (Newton, Alfred, 1861), cited in (Gaskell 2000), also see Messybeast: http://Messybeast.com/Extinct/Great-Auk.htm and (Gísli Pálsson. 2024) for discussion. The eye structure of penguins allows them to see both on land and underwater, which may be an example of convergent evolution seen on the Great Auk as well. ↩︎

  62. Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p155. ↩︎

  63. This is in French. ↩︎

  64. Baxter, James Phinney, Jean François De La Roque Roberval, and Jean Alfonse. A memoir of Jacques Cartier, sieur de Limoilou, his voyages to the St. Lawrence, a bibliography and a facsimile of the manuscript of. New York, Dodd, Mead & Co, 1906. Pdf. https://www.loc.gov/item/06013927/. p77-78. ↩︎

  65. Voyages and travels : ancient and modern, with introductions, notes and illustrations. New York : P. F. Collier and son, [c1910] The Harvard classics, ed. by C. W. Ellot [vol. XXXIII]. Original source: Sir Humphrey Gilbert’s Voyage to Newfoundland. Edward Haies. 1583. ↩︎

  66. Whitbourne R. A Discourse and Discouery of Nevv-Found-Land with Many Reasons to Prooue How Worthy and Beneficiall a Plantation May There Be Made, after a Far Better Manner than Now It Is. Together with the Laying Open of Certaine Enormities and Abuses Commited by Some That Trade to That Countrey, and the Meanes Laide Downe for Reformation Thereof. Written by Captaine Richard Whitbourne of Exmouth, in the County of Deuon, and Published by Authority[M]. Imprinted at London:: By Felix Kyngston, for William Barret, 1983. ↩︎

  67. This is in Latin, referring to the webbed feet. ↩︎

  68. This record is from before the 1752 calendar reform in Britain, so to convert it to the modern Gregorian calendar, add 11 days. See: Gaskell J. Who killed the great auk?[M]. Oxford University Press, 2000. p84. ↩︎

  69. Martin, M.1698. A late voyage to St. Kilda, the remotest of all the Hebrides, or the Western isles of Scotland with a history of the island, natural moral, and topographical : wherein is an account of their customes region, fish, fowl, &c. : as also a relation of a late imposter there, pretended to be sent by St. John Baptist / by M. Martin, gent. Oxford Test Archive, Oxford. ↩︎

  70. Old measurement unit: 1 league is equivalent to today’s 3 miles, approximately 4.8 kilometers. ↩︎

  71. Measurement unit in the liquor industry: one puncheon is about 70 to 120 gallons. ↩︎

  72. Thomas, Aaron. The Newfoundland Journal of Aaron Thomas, Able Seaman in H.M.S. Boston: A Journal Written During a Voyage from England to Newfoundland and from Newfoundland to England in the Years 1794 and 1795, Addressed to a Friend. United Kingdom: Longmans, 1968. For details on the slaughter of Great Auks, see p127. Quoted from: (Fuller, 1999), (Gaskell 2000). Mowat, Farley. Sea of slaughter. Boston: Atlantic Monthly Press, 1984. and Stark, Lizzie. Egg: A Dozen Ovatures. United States: W. W. Norton, 2023. ↩︎

  73. This implies that Great Auks could lay more than one egg per year, which contradicts the consensus. See the discussion in (Gaskell 2000). ↩︎

  74. The cruelty described in these accounts is so extreme that modern natural historians question their authenticity. Some believe the descriptions are “gory, obviously distorted and exaggerated,” but no specific reasons are given. As firsthand accounts, they are likely accurate. See Montevecchi, William A., and Leslie M. Tuck. “Newfoundland birds: exploitation, study, conservation.” (1987). p57, and the discussion in (Fuller, 1999). (Gaskell 2000) notes that Aaron Thomas’s tone when describing the killing of Great Auks is strangely flippant compared to the clear, solemn English in other parts of his Newfoundland diary, possibly reflecting a defensive psychological reaction to the cruelty. (Gaskell 2000) also discusses the practical reasons for burning Great Auk bodies in kettles, suggesting it was to remove feathers and eliminate parasites, but also cites less direct sources claiming sailors burned Great Auks as purely malicious mischief. ↩︎

  75. Fleming, J., 1823. Gleanings of natural history, gathered on the coast of Scotland during a voyage in 1821. Edinb. Phil. J. 8 : 294-303. This Great Auk later escaped with the rope still attached to its leg, and its whereabouts are unknown. ↩︎

  76. There were a total of 14 people, but only 3 went ashore. By 1858, 2 of them had died. John Wolley and Alfred Newton interviewed all the remaining 12 individuals. ↩︎

  77. Newton, Alfred. “XLII.—Abstract of Mr. J. Wolley’sResearches in Iceland respecting the Gare‐fowl or Great Auk (Alea impennis, Linn.).” Ibis 3, no. 4 (1861): 374-399. ↩︎

  78. The ages of the three men in 1844 were 25, 41, and 21 years respectively. See: The Gare-Fowl Books. John Wolley. Cambridge University Library. 转引自Gísli Pálsson. 2024. The Last of Its Kind. Princeton University Press. ↩︎

  79. Refers to the Black Guillemot (Cepphus grylle). ↩︎

  80. Old measurement unit, 1 fathom equals 6 feet, approximately 1.8288 meters. ↩︎

  81. The pronoun “him” is used throughout this recollection, but the sex of this Great Auk cannot be confirmed. According to Wolley’s interview notes, the other Great Auk was incubating an egg at the time, but both male and female Great Auks participated in incubation, so this is not a reliable basis for determining sex. ↩︎

  82. The Gare-Fowl Books. John Wolley. Cambridge University Library. Quoted in Fuller Errol. The Great Auk. 1999. p82. ↩︎ ↩︎ ↩︎

  83. According to Herr Brodtkorb, these birds had only stubs for wings and white spots near their eyes. These characteristics can identify the Great Auk, and Robert Collet vouched for Herr Brodtkorb’s ability to identify seabirds. However, the location remains a significant doubt. ↩︎

  84. Letter from Brodtkorb to Collet, 1883. Quoted in Fuller, Errol. The Great Auk. 1999. p408. ↩︎